Glasgow duo Jamie Dobie and Stafford McDowall – who both played more than half of the Ireland game after coming on for Graham and Russell – are again the two replacement backs as Gregor Townsend’s side head to Twickenham bidding to win the Calcutta Cup for the fifth year in succession.
There are two changes to the Scottish pack, with Edinburgh pair Pierre Schoeman and Jamie Ritchie taking over from Glasgow duo Rory Sutherland and Matt Fagerson in the front row and back row respectively.
Scotland co-captain Finn Russell has been passed fit for Saturday’s Calcutta Cup match with England but fellow back Darcy Graham misses out.
The key duo were doubts for the Guinness Six Nations showdown at Allianz Stadium after colliding with each other and being forced off with head injuries in the first half of the defeat by Ireland.
Sutherland and Fagerson both drop to the bench where they are joined by fellow forwards Ewan Ashman, Will Hurd, Sam Skinner and Gregor Brown in a 6-2 split.
